Navigator and his attendant the Man of Iron make up the crew of a small Autobot rescue ship. Little is known about him, but he is probably good at navigating. He seems to operate the ship with his own onboard systems.
Marvel Comics UK continuity
Navigator and the Man of Iron were sent by the Autobots (sometime between four hundred and four million years ago) to locate the missing Ark. They successfully located Earth, but since the Ark was offline, it did not return their rescue signal. Navigator patiently waited in the ship, underground, while the Man of Iron periodically made sure the surroundings were safe. Ultimately, when Optimus Prime, Megatron and their crews reawakened, a pitched battle was fought to recover Navigator's ship. The result was the destruction of the ship and Navigator himself at the hands of the unwitting Jazz.